Inhibiting immune checkpoint molecules – including programmed cell death protein (PD-1) and cytotoxic T-lymphocyte-associated-protein 4 (CTLA-4) – can induce T cell-mediated killing of cancer cells and results in significant clinical responses in ~20% of non-small-cell lung cancer patients. However, never-smokers with lung cancer, who currently comprise 20% of lung cancer cases in Australia, typically respond poorly to checkpoint immunotherapy. Literature suggests that tumour cell intrinsic factors – including lower PDL1 expression by tumour cells and lower tumour mutational burden – account for reduced sensitivity to checkpoint inhibition. Yet tumour cell extrinsic factors, including immune parameters, are yet to be formally explored. To this end, we profiled 10 never-smoker and 10 ever-smoker patient lung and matched lung tumours with mass cytometry to explore the effects of smoking on resident memory T cells (TRM) in the human lung, and how this impacts immune infiltrate in tumours. We found lower co-stimulatory marker expression on CD4+ TRM in never-smoker lung tissue and accordingly less CD8+ TRM and resident CD4+ T regulatory cells compared to ever-smokers. In tumour tissue, infiltrating CD4+ and CD8+ TRM had less co-stimulatory and co-inhibitory (including PD-1 and CTLA4) marker expression in never-smokers. This data suggests less T cell activation in the normal lung and resultant lung tumour in never-smokers with lung cancer. Tumour cells isolated from never-smokers expressed either MHC Class I and/or MHC Class II, whereas tumours that had lost both MHC molecules were only detected in ever-smokers – potentially reflecting an outcome of differing levels of immune-surveillance during tumour development. Collectively, our data suggests that a lack of initial T cell response to tumours is a contributing factor to the inherent resistance of never-smoker patients to anti-PD1 or anti-CTLA4 therapy. We are currently focused on instead targeting co-stimulatory molecules on tumour-infiltrating T cells in never-smoker lung cancer. Our study suggests that individualized strategies will be necessary for the successful treatment of ever- and never-smoker lung cancer with immunotherapies.